Dab the starter and the Tuono V4R’s 999.6cc V4 fires quickly, the tachometer jumping as the ECU automatically blips the ride-by-wire throttle. Tuono means “thunder” in Italian, and while that name might have better suited the original Mille-based V-twin, the current V4 still makes itself known with a raspy bark overlaying a deep, booming bass line. Updating the decade-old Tuono wasn’t as simple as stripping the bodywork from Aprilia’s RSV4 superbike; the engine, chassis and riding position all were altered to tailor the bike to the street. The 65-degree V4 produces 145 bhp at 11,800 rpm. That’s 7 bhp down on the world-beating RSV4, but this engine has been tuned for more midrange power with revised valve timing and longer velocity stacks. Performance is enhanced with a heavier flywheel and shortened gear ratios in first through third. Aprilia also lowered the swingarm pivot and the engine position to bring the center of gravity closer to the ground. Like the RSV4R APRC, the Tuono comes equipped with the Aprilia Performance Ride Control electronics package that includes eight-level traction control, variable power modes, a quick-shifter, wheelie control and launch control.